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An adventure/RPG based on the 3D animated action-adventure television series, Code Lyoko is about a group of students who uncover a parallel universe called Lyoko inside a super-computer that is threatened by a virus that could destroy the universe. Four ordinary students -- Jeremy, Odd, Ulrich and Yumi -- must help a virtual humanoid creature named "Aelita" to stop rogue computer XANA's latest attack on their real world.
Changing tactics, they realise they can use "Translation" to materialise in the labs that contain each supercomputer, which are hidden around the globe, protected by bosses.
Each end-level boss is a guardian that exists in the physical world, and has been built or enhanced. The heroes must confront all the bosses and destroy all the supercomputers before achieving their ultimate goal of saving William.
By scanning the Digital Sea, the heroes discover that the supercomputers sustaining the Replikas which they destroyed using the "Chimera" code are linked to a mysterious and dangerous Replika: the Volcano Replika.
It soon becomes clear that this Replika contains the true core of X.A.N.A: a crater containing a number of towers.
When they finally manage to destroy all the other supercomputers; they turn their entire efforts to the supercomputer in this Replika. They discover the tower controlling William in the crater on the Volcano Replika and manage to save him just before destroying the Replika, while destroying X.A.N.A at the same time!